TF JASPAR ID Class Family Tax Group IC GC Content Target gene hits Target gene non-hits Background gene hits Background gene non-hits Target TFBS hits Target TFBS nucleotide rate Background TFBS hits Background TFBS nucleotide rate Z-score Fisher score NFYA MA0060.1 Other Alpha-Helix NFY CCAAT-binding vertebrates 12.925 0.523 22 19 5865 18887 39 0.00838 9476 0.00421 17.524 10.288 Foxa2 MA0047.2 Winged Helix-Turn-Helix Forkhead vertebrates 13.268 0.315 26 15 11656 13096 123 0.0198 43535 0.0145 12.084 3.638 FOXF2 MA0030.1 Winged Helix-Turn-Helix Forkhead vertebrates 14.824 0.334 14 27 4097 20655 22 0.00414 6459 0.00251 8.811 5.355 SRY MA0084.1 Other Alpha-Helix High Mobility Group vertebrates 9.193 0.238 33 8 14714 10038 249 0.0301 101929 0.0255 7.955 5.612 FOXA1 MA0148.1 Winged Helix-Turn-Helix Forkhead vertebrates 12.533 0.332 29 12 13355 11397 157 0.0232 63112 0.0193 7.723 3.837 RORA_2 MA0072.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 17.425 0.369 13 28 3571 21181 17 0.0032 5079 0.00198 7.456 5.540 FOXO3 MA0157.1 Winged Helix-Turn-Helix Forkhead vertebrates 11.734 0.298 32 9 13611 11141 175 0.0188 69410 0.0154 7.446 6.262 Sox2 MA0143.1 Other Alpha-Helix High Mobility Group vertebrates 12.951 0.361 9 32 2809 21943 13 0.00262 3844 0.0016 6.888 3.276 ARID3A MA0151.1 Helix-Turn-Helix Arid vertebrates 9.896 0.062 36 5 16086 8666 393 0.0317 166235 0.0277 6.552 6.958 Prrx2 MA0075.1 Helix-Turn-Helix Homeo vertebrates 9.063 0.027 33 8 15063 9689 296 0.0199 123270 0.0171 5.763 5.096 FOXD1 MA0031.1 Winged Helix-Turn-Helix Forkhead vertebrates 11.926 0.275 32 9 13087 11665 153 0.0164 62921 0.014 5.675 7.137 Nkx2-5 MA0063.1 Helix-Turn-Helix Homeo vertebrates 8.270 0.218 35 6 16973 7779 448 0.0421 197210 0.0384 5.320 4.420 Lhx3 MA0135.1 Helix-Turn-Helix Homeo vertebrates 16.354 0.131 14 27 5806 18946 34 0.00594 12758 0.00461 5.318 2.522 NKX3-1 MA0124.1 Helix-Turn-Helix Homeo vertebrates 11.127 0.179 31 10 12654 12098 165 0.0155 68823 0.0134 5.024 6.740 NFATC2 MA0152.1 Ig-fold Rel vertebrates 9.859 0.346 34 7 15176 9576 229 0.0215 97757 0.019 5.003 5.992 Pax6 MA0069.1 Helix-Turn-Helix Homeo vertebrates 13.798 0.432 4 37 1218 23534 5 0.00094 1377 0.000536 4.689 1.953 HOXA5 MA0158.1 Helix-Turn-Helix Homeo vertebrates 8.759 0.315 37 4 17629 7123 458 0.0492 205993 0.0458 4.441 5.732 E2F1 MA0024.1 Winged Helix-Turn-Helix E2F vertebrates 13.838 0.625 17 24 8113 16639 46 0.00494 17630 0.00392 4.436 1.868 Foxd3 MA0041.1 Winged Helix-Turn-Helix Forkhead vertebrates 12.945 0.223 29 12 11354 13398 115 0.0185 49535 0.0165 4.297 6.778 HNF1B MA0153.1 Helix-Turn-Helix Homeo vertebrates 16.821 0.222 12 29 4107 20645 19 0.00306 6910 0.0023 4.273 3.483 MEF2A MA0052.1 Other Alpha-Helix MADS vertebrates 15.709 0.179 17 24 7006 17746 40 0.00537 15631 0.00434 4.237 3.025 TLX1::NFIC MA0119.1 Helix-Turn-Helix::Other Homeo::Nuclear Factor I-CCAAT-binding vertebrates 19.665 0.598 4 37 1066 23686 4 0.000752 1162 0.000452 3.764 2.309 FOXI1 MA0042.1 Winged Helix-Turn-Helix Forkhead vertebrates 13.183 0.263 31 10 11680 13072 113 0.0182 49366 0.0165 3.733 8.506 Pdx1 MA0132.1 Helix-Turn-Helix Homeo vertebrates 9.040 0.194 34 7 15507 9245 317 0.0255 140973 0.0235 3.657 5.483 CEBPA MA0102.2 Zipper-Type Leucine Zipper vertebrates 8.712 0.358 28 13 12828 11924 128 0.0155 55606 0.0139 3.626 3.711 Esrrb MA0141.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 12.806 0.524 27 14 10167 14585 62 0.00999 26346 0.00878 3.506 6.730 RORA_1 MA0071.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 13.190 0.424 24 17 8624 16128 46 0.00618 18895 0.00525 3.473 6.407 HLF MA0043.1 Zipper-Type Leucine Zipper vertebrates 11.147 0.394 15 26 5508 19244 25 0.00403 9969 0.00332 3.307 3.634 Nr2e3 MA0164.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 12.028 0.373 18 23 6354 18398 31 0.00291 12287 0.00239 2.891 4.757 TAL1::TCF3 MA0091.1 Zipper-Type Helix-Loop-Helix vertebrates 14.070 0.453 18 23 7491 17261 35 0.00564 14681 0.0049 2.884 3.100 NR2F1 MA0017.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 15.924 0.478 12 29 3735 21017 13 0.00244 5073 0.00197 2.851 4.171 TBP MA0108.2 Beta-sheet TATA-binding vertebrates 10.086 0.377 22 19 10389 14363 76 0.0153 33775 0.0141 2.834 2.428 NHLH1 MA0048.1 Zipper-Type Helix-Loop-Helix vertebrates 14.132 0.674 14 27 5783 18969 24 0.00387 9874 0.00329 2.706 2.549 NFKB1 MA0105.1 Ig-fold Rel vertebrates 15.627 0.758 12 29 3946 20806 16 0.00236 6372 0.00195 2.531 3.767 ELK4 MA0076.1 Winged Helix-Turn-Helix Ets vertebrates 14.123 0.583 19 22 6034 18718 23 0.00278 9342 0.00234 2.468 6.318 NFIL3 MA0025.1 Zipper-Type Leucine Zipper vertebrates 14.139 0.265 19 22 6731 18021 37 0.00547 15828 0.00484 2.440 4.999 Arnt MA0004.1 Zipper-Type Helix-Loop-Helix vertebrates 10.992 0.642 20 21 7052 17700 34 0.00274 13904 0.00232 2.352 5.352 SP1 MA0079.2 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 11.129 0.820 30 11 12391 12361 174 0.0234 79797 0.0222 2.196 6.105 Nobox MA0125.1 Helix-Turn-Helix Homeo vertebrates 9.573 0.260 29 12 13558 11194 169 0.0182 77190 0.0172 2.078 3.592 SRF MA0083.1 Other Alpha-Helix MADS vertebrates 17.965 0.466 3 38 908 23844 3 0.000483 1019 0.00034 2.027 1.659 Tcfcp2l1 MA0145.1 Other CP2 vertebrates 11.650 0.609 25 16 10591 14161 71 0.0133 32190 0.0125 2.008 4.230 MAX MA0058.1 Zipper-Type Helix-Loop-Helix vertebrates 12.685 0.571 18 23 7993 16759 36 0.00483 15900 0.00442 1.684 2.527 HIF1A::ARNT MA0259.1 Zipper-Type Helix-Loop-Helix vertebrates 9.740 0.657 26 15 10914 13838 83 0.00892 37662 0.00837 1.609 4.604 Evi1 MA0029.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 17.909 0.280 5 36 1931 22821 6 0.00113 2457 0.000956 1.461 1.544 PLAG1 MA0163.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 19.352 0.798 5 36 1971 22781 6 0.00113 2462 0.000958 1.442 1.489 Pou5f1 MA0142.1 Helix-Turn-Helix Homeo vertebrates 14.808 0.308 9 32 3402 21350 12 0.00242 5205 0.00217 1.413 2.285 Egr1 MA0162.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 14.456 0.739 13 28 6364 18388 27 0.00399 12188 0.00373 1.147 1.436 CTCF MA0139.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 17.205 0.645 8 33 3236 21516 9 0.0023 3982 0.0021 1.115 1.834 MIZF MA0131.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 13.197 0.610 6 35 2080 22672 6 0.000806 2497 0.000694 1.089 2.067 IRF1 MA0050.1 Winged Helix-Turn-Helix IRF vertebrates 16.008 0.383 16 25 6075 18677 25 0.00403 11517 0.00384 0.800 3.559 Klf4 MA0039.2 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 12.618 0.771 32 9 13645 11107 187 0.0251 88731 0.0247 0.788 6.207 SPIB MA0081.1 Winged Helix-Turn-Helix Ets vertebrates 9.060 0.466 36 5 17547 7205 369 0.0347 175639 0.0342 0.773 4.665 NR3C1 MA0113.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 14.749 0.432 6 35 2522 22230 7 0.00169 3161 0.00158 0.716 1.444 Tal1::Gata1 MA0140.1 Zipper-Type Helix-Loop-Helix vertebrates 11.297 0.451 9 32 6212 18540 24 0.0058 11200 0.0056 0.703 0.309 Stat3 MA0144.1 Ig-fold Stat vertebrates 13.601 0.505 24 17 9883 14869 50 0.00671 23645 0.00657 0.463 4.396 NF-kappaB MA0061.1 Ig-fold Rel vertebrates 13.345 0.621 21 20 8381 16371 40 0.00537 18865 0.00524 0.463 4.112 Arnt::Ahr MA0006.1 Zipper-Type Helix-Loop-Helix vertebrates 9.532 0.715 28 13 13597 11155 139 0.0112 66086 0.011 0.455 2.850 Pax5 MA0014.1 Helix-Turn-Helix Homeo vertebrates 12.432 0.575 2 39 1224 23528 3 0.000806 1364 0.000758 0.406 0.497 IRF2 MA0051.1 Winged Helix-Turn-Helix IRF vertebrates 21.134 0.412 2 39 844 23908 2 0.000483 923 0.000462 0.191 0.891 USF1 MA0093.1 Zipper-Type Helix-Loop-Helix vertebrates 11.290 0.643 25 16 9330 15422 45 0.00423 21993 0.00428 -0.229 6.157 NR4A2 MA0160.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 10.165 0.491 36 5 13896 10856 123 0.0132 60034 0.0133 -0.330 11.181 MYC::MAX MA0059.1 Zipper-Type Helix-Loop-Helix vertebrates 14.237 0.602 8 33 3436 21316 9 0.00133 4535 0.00139 -0.466 1.601 Ar MA0007.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 15.703 0.498 1 40 534 24218 1 0.000295 554 0.000339 -0.741 0.525 RELA MA0107.1 Ig-fold Rel vertebrates 14.757 0.567 17 24 7171 17581 27 0.00363 13690 0.0038 -0.821 2.825 Sox5 MA0087.1 Other Alpha-Helix High Mobility Group vertebrates 10.831 0.199 23 18 13087 11665 145 0.0136 72042 0.014 -0.906 0.916 Ddit3::Cebpa MA0019.1 Zipper-Type Leucine Zipper vertebrates 11.652 0.483 14 27 5576 19176 19 0.00306 9754 0.00325 -0.947 2.807 ESR2 MA0258.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 13.618 0.546 4 37 2219 22533 5 0.00121 2670 0.00134 -0.999 0.678 AP1 MA0099.2 Zipper-Type Leucine Zipper vertebrates 9.193 0.405 34 7 16390 8362 227 0.0213 113253 0.022 -1.299 4.233 TP53 MA0106.1 Zinc-coordinating Loop-Sheet-Helix vertebrates 26.239 0.603 0 41 8 24744 0 0 8 4.45e-06 -1.444 0.000 PBX1 MA0070.1 Helix-Turn-Helix Homeo vertebrates 14.641 0.310 8 33 4107 20645 12 0.00193 6634 0.00221 -1.656 0.997 Foxq1 MA0040.1 Winged Helix-Turn-Helix Forkhead vertebrates 14.070 0.202 18 23 8007 16745 42 0.0062 22088 0.00675 -1.847 2.512 REL MA0101.1 Ig-fold Rel vertebrates 10.515 0.559 25 16 10514 14238 56 0.00752 29250 0.00813 -1.869 4.334 Myb MA0100.1 Helix-Turn-Helix Myb vertebrates 9.883 0.591 33 8 13518 11234 114 0.0122 58600 0.013 -1.894 7.583 ELK1 MA0028.1 Winged Helix-Turn-Helix Ets vertebrates 8.812 0.568 30 11 13642 11110 107 0.0144 54748 0.0152 -1.899 4.287 PPARG MA0066.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 20.365 0.523 0 41 62 24690 0 0 64 3.56e-05 -1.935 0.000 MZF1_5-13 MA0057.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 9.400 0.588 30 11 13425 11327 153 0.0205 77649 0.0216 -1.950 4.576 SPI1 MA0080.2 Winged Helix-Turn-Helix Ets vertebrates 9.640 0.435 33 8 15792 8960 188 0.0177 95792 0.0186 -1.953 4.099 NFE2L2 MA0150.1 Zipper-Type Leucine Zipper vertebrates 14.394 0.486 8 33 5635 19117 16 0.00236 8970 0.00274 -2.010 0.292 NR1H2::RXRA MA0115.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 27.878 0.445 0 41 99 24653 0 0 100 4.72e-05 -2.142 0.000 T MA0009.1 Beta-Hairpin-Ribbon T vertebrates 17.863 0.452 3 38 1811 22941 3 0.000443 2114 0.000646 -2.253 0.535 HNF4A MA0114.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 9.617 0.522 16 25 6389 18363 20 0.00349 11240 0.00406 -2.471 3.125 SOX9 MA0077.1 Other Alpha-Helix High Mobility Group vertebrates 9.079 0.358 26 15 11417 13335 78 0.00943 41403 0.0104 -2.519 3.933 Gfi MA0038.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 9.470 0.391 29 12 13731 11021 127 0.0171 65844 0.0183 -2.544 3.391 Pax4 MA0068.1 Helix-Turn-Helix Homeo vertebrates 11.004 0.441 0 41 112 24640 0 0 113 9.42e-05 -2.838 0.000 ELF5 MA0136.1 Winged Helix-Turn-Helix Ets vertebrates 8.693 0.429 35 6 16547 8205 228 0.0276 117435 0.0294 -2.943 5.023 EWSR1-FLI1 MA0149.1 Winged Helix-Turn-Helix Ets vertebrates 32.871 0.554 0 41 207 24545 0 0 209 0.000105 -2.969 0.000 FEV MA0156.1 Winged Helix-Turn-Helix Ets vertebrates 12.121 0.442 34 7 15402 9350 155 0.0167 81722 0.0182 -3.111 5.642 RUNX1 MA0002.2 Ig-fold Runt vertebrates 10.401 0.502 30 11 13641 11111 111 0.0164 58648 0.0179 -3.162 4.288 GABPA MA0062.2 Winged Helix-Turn-Helix Ets vertebrates 13.335 0.647 21 20 9355 15397 37 0.00547 20927 0.0064 -3.211 2.894 REST MA0138.2 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 23.134 0.596 0 41 232 24520 0 0 242 0.000141 -3.397 0.000 Myc MA0147.1 Zipper-Type Helix-Loop-Helix vertebrates 11.157 0.686 17 24 8288 16464 31 0.00416 18476 0.00513 -3.734 1.720 HNF1A MA0046.1 Helix-Turn-Helix Homeo vertebrates 15.548 0.259 5 36 3044 21708 6 0.00113 4352 0.00169 -3.795 0.546 Nkx3-2 MA0122.1 Helix-Turn-Helix Homeo vertebrates 8.542 0.463 34 7 15425 9327 172 0.0208 91612 0.0229 -3.887 5.607 ZNF354C MA0130.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 8.958 0.615 38 3 17566 7186 397 0.032 207738 0.0346 -3.964 7.267 ESR1 MA0112.2 Zinc-coordinating Hormone-nuclear Receptor vertebrates 13.563 0.594 0 41 345 24407 0 0 355 0.000197 -3.964 0.000 RXRA::VDR MA0074.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 20.451 0.527 0 41 453 24299 0 0 475 0.000198 -3.970 0.000 Mycn MA0104.2 Zipper-Type Helix-Loop-Helix vertebrates 11.104 0.699 17 24 8332 16420 31 0.00416 18928 0.00526 -4.164 1.684 Gata1 MA0035.2 Zinc-coordinating GATA vertebrates 10.878 0.373 30 11 14350 10402 141 0.0208 75905 0.0232 -4.316 3.417 TEAD1 MA0090.1 Helix-Turn-Helix Homeo vertebrates 15.678 0.507 10 31 5222 19530 12 0.00193 8446 0.00282 -4.580 1.020 MZF1_1-4 MA0056.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 8.586 0.725 36 5 16863 7889 378 0.0305 201000 0.0335 -4.645 5.688 YY1 MA0095.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 8.101 0.510 34 7 17190 7562 307 0.0247 165054 0.0275 -4.655 3.237 EBF1 MA0154.1 Zipper-Type Helix-Loop-Helix vertebrates 11.564 0.648 28 13 11288 13464 67 0.009 39149 0.0109 -4.968 5.861 RXR::RAR_DR5 MA0159.1 Zinc-coordinating Hormone-nuclear Receptor vertebrates 16.004 0.535 1 40 1313 23439 1 0.000228 1499 0.000708 -4.991 0.113 CREB1 MA0018.2 Zipper-Type Leucine Zipper vertebrates 10.139 0.523 17 24 9204 15548 29 0.00312 19712 0.00438 -5.261 1.082 Sox17 MA0078.1 Other Alpha-Helix High Mobility Group vertebrates 10.502 0.384 30 11 13879 10873 116 0.014 65883 0.0165 -5.283 3.983 znf143 MA0088.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 17.541 0.530 1 40 1301 23451 1 0.000269 1473 0.000819 -5.313 0.116 Hand1::Tcfe2a MA0092.1 Zipper-Type Helix-Loop-Helix vertebrates 10.144 0.507 27 14 13286 11466 95 0.0128 54768 0.0152 -5.503 2.539 Zfx MA0146.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 13.077 0.749 25 16 9872 14880 56 0.0105 33238 0.0129 -5.820 5.269 INSM1 MA0155.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 14.862 0.667 15 26 7588 17164 26 0.00419 17685 0.0059 -6.110 1.374 RREB1 MA0073.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 22.278 0.623 1 40 1520 23232 1 0.000269 1839 0.00102 -6.492 0.077 Spz1 MA0111.1 Other Other vertebrates 11.907 0.538 14 27 6658 18094 16 0.00236 12530 0.00383 -6.509 1.658 Zfp423 MA0116.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 17.925 0.679 10 31 5691 19061 12 0.00242 10149 0.00423 -7.652 0.743 Myf MA0055.1 Zipper-Type Helix-Loop-Helix vertebrates 15.914 0.609 18 23 10157 14595 40 0.00645 27519 0.00918 -7.834 0.887 STAT1 MA0137.2 Ig-fold Stat vertebrates 13.119 0.452 9 32 6394 18358 13 0.00262 10990 0.00458 -7.957 0.264 PPARG::RXRA MA0065.2 Zinc-coordinating Hormone-nuclear Receptor vertebrates 11.663 0.547 9 32 6067 18685 13 0.00262 11080 0.00462 -8.076 0.349 ZEB1 MA0103.1 Zinc-coordinating BetaBetaAlpha-zinc finger vertebrates 8.305 0.557 36 5 17454 7298 292 0.0235 171132 0.0285 -8.216 4.799